Ph.D. in Plant sciences
Shanghai Jiao Tong University, Shanghai, China

Master of Science in Botany
Abdul Wali Khan University Mardan, Pakistan

General Biology (with Laboratory)

Genetics (with Laboratory)

Research and Technology

Dr. Muhammad Khalid, received his master’s degree from Abdul Wali Khan University Mardan, Pakistan. Subsequently, he obtained his doctoral degree from Shanghai Jiao Tong University, China, where his research focus was directed towards the exploration of plant-microbe interactions. Dr. Khalid continued his research as a postdoctoral fellow at the same university, delving into the realm of environmental ecology. During his postdoctoral fellowship, he collaborated with the University of Helsinki, Finland, to complete several national and international projects. He was awarded Shanghai Jiao Tong University technical assistant researcher award in 2021 and the Morning Star program award in 2022 based on outstanding research performance.

Currently, Dr. Khalid is imparting his knowledge and expertise as a biology lecturer at Wenzhou-Kean University in China, where he teaches courses such as general biology, genetics, and research and technology.

Dr. Khalid, enjoys playing football and badminton as hobbies. He has been an active member of the International Student Badminton Association of Shanghai Jiao Tong University.

Dr. Khalid’s research focus is on environmental microbial ecology, studying the microbiome’s association with plants and their soil environment. Additionally, he is passionate about discovering new insights related to microbes-assisted phytoremediation.

1. Khalid, Muhammad, Haoxin Tan, Mehran Ali, Asad Rehman, Xinxin Liu, Lantian Su, Chang Zhao, Xiaoxiao Li, and Nan Hui. “Karst rocky desertification diverged the soil residing and the active ectomycorrhizal fungal communities thereby fostering distinctive extramatrical mycelia.” Science of The Total Environment 807 (2022): 151016. https://doi.org/10.1016/j.scitotenv.2021.151016
2. Khalid, Muhammad, Baoming Du, Haoxin Tan, Xinxin Liu, Lantian Su, Mehran Ali, Chunjiang Liu, Ningxiao Sun, and Nan Hui. “Phosphorus elevation erodes ectomycorrhizal community diversity and induces divergence of saprophytic community composition between vegetation types.” Science of The Total Environment (2021): 148502. https://doi.org/10.1016/j.scitotenv.2021.148502
3. Khalid, Muhammad, Mehran Ali, Danial Hassani, Abdur Rauf, Farooq Jan, and Nan Hui. “Salicylic acid mediated protection of Brassica campestris sp. chinensis from saline stress via SA receptor NPR1 dependent transcriptional regulation and biosynthesis of related biochemicals.” Environmental Technology & Innovation 24 (2021): 101950. https://doi.org/10.1016/j.eti.2021.101950
4. Khalid, Muhammad, UR-RAHMAN Saeed, Danial Hassani, Kashif Hayat, Z. H. O. U. Pei, and H. U. I. Nan. “Advances in fungal-assisted phytoremediation of heavy metals: A review.” Pedosphere 31, no. 3 (2021): 475-495. https://doi.org/10.1016/s1002-0160(20)60091-1
5. Khalid, Muhammad, Saeed-ur Rahman, and Danfeng Huang. “Molecular mechanism underlying Piriformospora indica-mediated plant improvement/protection for sustainable agriculture.” Acta biochimica et biophysica Sinica 51, no. 3 (2019): 229-242. https://doi.org/10.1093/abbs/gmz004
6. Khalid, Muhammad, UR-RAHMAN Saeed, T. A. N. Haoxin, S. U. Lantian, Z. H. O. U. Pei, and H. U. I. Nan. “Mutualistic fungus Piriformospora indica modulates cadmium phytoremediation properties of host plant via concerted action of enzymatic and non-enzymatic biochemicals.” Pedosphere 32, no. 2 (2022): 256-267. https://doi.org/10.1016/s1002-0160(21)60014-0
7. Khalid, Muhammad, Danial Hassani, Jianli Liao, Xin Xiong, Muhammad Bilal, and Danfeng Huang. “An endosymbiont Piriformospora indica reduces adverse effects of salinity by regulating cation transporter genes, phytohormones, and antioxidants in Brassica campestris ssp. Chinensis.” Environmental and experimental botany 153 (2018): 89-99. https://doi.org/10.1016/j.envexpbot.2018.05.007
8. Hassani, Danial, Xueqing Fu, Qian Shen, Muhammad Khalid, Jocelyn KC Rose, and Kexuan Tang. “Parallel transcriptional regulation of artemisinin and flavonoid biosynthesis.” Trends in plant science 25, no. 5 (2020): 466-476. https://doi.org/10.1016/j.tplants.2020.01.001
9. Khalid, Muhammad, Muhammad Bilal, and Dan-feng HUANG. “Role of flavonoids in plant interactions with the environment and against human pathogens—A review.” Journal of integrative agriculture 18, no. 1 (2019): 211-230. https://doi.org/10.1016/S2095-3119(19)62555-4
10. Khalid, Muhammad, Danial Hassani, Muhammad Bilal, Jianli Liao, and Danfeng Huang. “Elevation of secondary metabolites synthesis in Brassica campestris ssp. chinensis L. via exogenous inoculation of Piriformospora indica with appropriate fertilizer.” PloS one 12, no. 5 (2017): e0177185. https://doi.org/10.1371/journal.pone.0177185
11. Khalid, Muhammad, Danial Hassani, Muhammad Bilal, Zahid Ali Butt, Muhammad Hamayun, Ayaz Ahmad, Danfeng Huang, and Anwar Hussain. “Identification of oral cavity biofilm forming bacteria and determination of their growth inhibition by Acacia arabica, Tamarix aphylla L. and Melia azedarach L. medicinal plants.” Archives of oral biology 81 (2017): 175-185. https://doi.org/10.1016/j.archoralbio.2017.05.011